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alternating whorls Ananiev Andrews l960 arrow articulation attachment of stalk basal bearing two sporangia Belg bifurcated Bowmanites bracts branching pattern Calamitaceae Calamophyton bicephalum Calamophyton primaevum Calamophyton renieri Carboniferous character complex composed cone described Devonian flora diagnosis of Calamophyton digitate branching distal elongate Equisetales erect appendage Eviostachya fertile axes forbesii forked genera genus Geol Givetian H^eg Haspia Hjrfeg Hyenia elegans Hyenia sphenophylloides intercalated Krausel and Weyland larger axes lateral recurved lateral stalk leafy Leclercq and Andrews long sterile lower main axis Middle Devonian morphology Nathorst Neiburg number of dichotomies ornamentation pairs phores Pl.l portion Protoarticulatae protoarticulates Pseudosporochnus Reconstruction recurved laterals recurved sporangial stalks revealed rhizome segment showed the attachment specialized spike species Sphenophyllales sphenopsids spinae sporangial arms sporangiophore axis sporangium attached spores sporophyll sterile appendage sterile leaves sterile projection superposed terminal text fig text figure three lateral tracheids trilete mark twelve sporangia uncovering verticillate vogtii whorls of six xylem