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acetylene reduction activity acetylene reduction rates algal blooms algal nitrogen ammonium Anabaena Aphanizomenon flos-aquae Arbor Vitae Lake August Big Arbor Vitae blue-green algae C2H2 reduction CM CM CM combined nitrogen culture decreased Deep Point depleted detached soybean nodules ethylene excised soybean nodules figure filaments filtered Fogg four replicates gas phase Gloeotrichia Green Bay growth heterocysts incubated July Lake Erie Lake Mendota Lake Michigan Lake Monona lake water Little Arbor Vitae maximum measured Microcystis N2-fixation N2-fixing nitrate nitrogen fixation nitrogenase nitrogenase activity nmoles C2H4 Nostoc muscorum Number oligotrophic phosphate phosphorus photosynthesis phytoplankton plankton rates of acetylene rates of C2H2 reduce acetylene reduction in Lake reduction per unit sampling stations Sanctuary Lake showed southern Green Bay species Station sampled stopper surface waters Table total nitrogen Trout Lake unit volume University Bay variation water samples Wisconsin lakes