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abundance adults allow analysis approximation artisanal fisheries artisanal fishing assessment average aztecus Berry Beverton and Holt biological biomass boat brown shrimp by-catch Caillouet calculated catchability caught cohort Costello curve cycle depends difficult economic rent equation estimate estuaries example exploitation rate factors FAO Fish.Rep fishermen fishing effort fishing mortality Garcia gear growth Gulf of Mexico Gulland increase intervals Ivory Coast juveniles lagoon larval length Lhomme Madagascar Marcille Maroochydore maximum maximum sustainable yield mesh Metapenaeus method migration moulting natural mortality nets observed obtained parameters penaeid shrimps Penaeus duorarum Penaeus notialis Penn period plankton population possible post-larvae prawn production recapture regression relation relationship rhythm Ricker salinity sampling seasonal variations Senegal setiferus showed shrimp fishery shrimp stocks size frequency distribution spawning species statistical tagged shrimps techniques trawl variability yield per recruit zones