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allele amplified polymorphic DNA arbitrary primers band patterns Beta vulgaris Brassica napus breeding lines bulked segregant analysis cloning clustering codominant combined competition correlation coupling phase detected DNA fragments DNA mixture DNA sequences genetic linkage map genome genotypes genotyping errors Hallden haploid heteroduplex heterozygous Hjerdin homozygous identified individuals large number linkage groups linkage phase mapping population marker-assisted selection markers in coupling molecular markers nematode resistance number of bands number of markers number of polymorphic phenotype Plant Breed polymorphic bands pools primers produced proportion random amplified polymorphic RAPD analysis RAPD and RFLP RAPD assay RAPD bands RAPD marker system RAPD markers RAPD markers linked RAPD primers RAPD protocol RAPD reactions RAPD system recombination region reproducibility resistance gene restorer allele restorer locus restriction enzymes RFLP and RAPD RFLP map RFLP markers segregating population set of primers single-primer specific subclustering sugar beet template DNA Theor Appl Genet two-primer mixtures