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aggrecan amplitude applied shear strain articular cartilage axial compression Axial LVDT axial modulus bath concentration behavior of cartilage biosynthetic response cartilage metabolism cartilage specimens Cartilage under Shear collagen compressive offset core and ring dermal punch diameter dynamic compression dynamic shear chamber Dynamic shear modulus dynamically sheared disks extracellular matrix Figure fluid flow free swelling control hyaluronan hydrostatic pressure incorporation in dynamically incorporation rate increasing ionic strength incubator induced linear regression Linear Stepper Motor Load Cell machine matrix shear mechanical behavior modulus behavior modulus under different modulus was measured NaCl outer ring region phase difference platens poroelastic proline proteoglycan pure shear mode relative fluid motion response of chondrocytes Ring of dynamically Rotary Positioning Table Rotational LVDT Holder samples Shear Application shear deformation shear modulus shear properties shear stress simple shear mode sinusoidal shear strain static control streaming potential stress relaxation sulfate Top Chamber Torque Cell torsional compliance viscoelastic