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acacia African species allopatric allospecies ally Angola appears areas arid barbets barred beak Bee-eater behaviour Benson bill birds breast breeding brown Bull Cameroun closely related coast colonies colour confined congeners Congo forest conspecific dark differences distinct distribution Eagle east eastern ecological endemic Ethiopia evergreen forest extends feathers feeding forms francolins genera genus geographical variation grasslands grey Guinea habitat highlands Ibis inhabits insects isolated Kenya known lowland Madagascar mainly migrant migratory monotypic montane nests Nigeria nightjars northern occurs overlap Palaearctic pale plumage populations probably race range records rectrices relationship rufous rump savanna season sedentary senegalensis similar smaller Somalia South Africa southern Africa species-group streaked subspecies Sudan suggests superspecies sympatric tail Tanzania three species throat treated trees tropical Uganda underparts upper West Africa western widely sympatric widespread wings wood-hoopoes woodland Woodpecker yellow Zaire Zambia