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adult air temperature animals anorexia anorexic antibiotic areas Atlas Mountains back legs basking facilities basking lamp become behaviour blood urea breeding calcium carbonate captivity carapace cause claws climate correct cure cuttlebone damage death diet differences disease effect egg shell faeces feeding female fertile flagellates Galapagos Giant Tortoise Greek Tortoise grow growth hatchlings healthy tortoise heat hence hibernate tortoises hibernation human humidity husbandry immune system incubation infection instincts keep the tortoise kidneys length lethal males mammals mating Mediterranean tortoises method mouth necessary nest northern Europe patterns Photo plastron possible problems protein ratio realise scute sexual sick tortoises smell soil sometimes species steadily subspecies summer tempera territory Testudo graeca Testudo hermanni Tethys Sea thermal tion toises tortoise appears tortoise normally tortoise's body temperature treatment ture urates urea uric acid urine veterinary vitamin warm waste products weight wild winter