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activated amino acid Andersson anti anti-CII antibody apoptosis arthritis in mice Arthritis Rheum arthritis susceptibility arthritogenic assay autoimmune disease autoreactive B-cell response binding bovine CII carbohydrates cartilage cartilage-specific chain chronic CII immunization class II molecules clone collagen type collagen-induced arthritis cytokines DBA/1 mice Del1 mice dendritic cells develop arthritis ELISPOT Engstrom epitope gene glycosylated H-2q mice heterologous Holmdahl human CII hybridomas hydroxylysine IFN-y immune response immune system immunodominant Immunology induced arthritis inflammatory interactions Jansson Klareskog lymph node lymphocytes major histocompatibility complex Malmstrom Metsaranta MHC class mice developed Michaelsson MMC-1 mice mouse CII mutated Natl Acad Sci nontransgenic littermates peptide peripheral tolerance Proc Natl Acad protein rat CII reactive receptor restricted rheumatoid arthritis sequence serum specific stimulation susceptible to CIA synovial T-cell T-cell epitope thymectomized thymectomy thymus tissue transgenic mice expressing TSC mice type II collagen Vuorio