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0:N ratio 1997 Sampling Date 30mm test diameter Adelaide Island AFDM ammonia excretion ammonium Antarctica ash free dry austral summer austral winter benthic benthos biochemical bivalve Brey Brockington calculated Chapter 4a chlorophyll chlorophyll standing stock Clarke coelomic Data are means droebachiensis Ebert Echinodermata echinoderms Echinoidea echinoids ectotherms elliptica energetic energy assimilated energy budget estimated Exp Mar Biol feeding period food availability free dry mass Giese gonad mass gonad tissue growth efficiency growth rate gut tissue increase indicate invertebrates latitude lipid Mar Biol Ecol marine invertebrates McMurdo measured month North Cove nutritional oocytes oxygen consumption rate Pearse phytoplankton polar conditions protein proximate composition range recorded reduced regressions reproductive output respiration Rothera Point Ryder Bay Sampling Date 1998 sea urchins seasonal energy seawater seawater temperature sediment shell length Signy Island size frequency distributions somatic growth South Cove spawning standard animal standard error Sterechinus neumayeri Strongylocentrotus values variation wild population