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20 March 9 April adult and juvenile adult cohort adult female surviving Alberta annual adult survival born per adult breeding season Buehler and Keith calculated central Wisconsin snowshoes composition of snowshoe corpora lutea date of first-litter days of recognizable decline Keith demographic development of snowshoe Estimated mean number eye lens weights Green and Evans habitat hare numbers hares in central Jun-Aug juvenile survival Keith and Windberg Kubisiak KUVLESKY L. B. Keith Lake Alexander lens weights Fig litter groups litters per adult litters per female mean conception dates mean date mortality northern Wisconsin number of days number of litters onset physical characteristics Keith population declines potential natality Prenatal Prenatal development realized natality recognizable pregnancy reproduction snow cover Snowshoe Hare distribution snowshoe hare populations species summer survival rates surviving the breeding testis weights thesis Tomahawk Total young born trap-night indices University of Wisconsin Wildl Wildlife Ecology Wisconsin snowshoe hares yearlings