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Abernethy abundance adult Alps August autumn behaviour Bendell black grouse black grouse Lyrurus black grouse population Black-billed Capercaillie blue grouse breeding brood caper Capercaillie and Black capercaillie Tetrao urogallus census Cervieres changes chicks clutch cocks counts decline display distribution ecology eggs Ellison factors females Finland Finnish fluctuations forest grouse forestry Galliformes grey partridge ground grouse Lyrurus tetrix Gullion habitat Hazel Grouse hens Hjorth hunting important juvenile Kinveachy Lagopus larch Lyrurus tetrix mainly males Manage Marcstrom mortality mountain myrtillus nesting success northern Norway number of capercaillie nutrition observed parvirostris pine Pinus planted forests population dynamics Porkert predation Pulliainen Rajala range red grouse region reproduction ruffed grouse Scotland Scots pine season Seiskari sex ratio shooting shot small-rodent snow species spruce grouse study area survival rate Sweden Table telemetry territory Tetrao urogallus tetraonids transects trees Vaccinium vegetation Wildl willow grouse winter woodland grouse young Zwickel