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abundant aerial surveys aircraft Alaska Department alaxensis aquatic plants arbusculoides arctic Alaska Average Berg and Phillips Betula nana Brooks Range browsed shrubs Coady collared moose Colville River daily movements Department of Fish deVos distance dwarf birch factor feeding Finland four habitat types Grizzly bears herbaceous plants histogram individual moose insects John Coady Killik River Kistchinski Loisa and Pulliainen long term observations magnitude of movements Minnesota moose activity moose habitat moose movements moose observed moose Radio moose were observed movement patterns movements of moose North NPR-A Number of moose occurred offriver open areas patterns of moose pellet groups preferred browse Pulliainen l968 relocations of collared Relocations of moose result river flood plain rutting Salix Salix alaxensis seasonal home ranges Seasonal Movement shrub areas shrubby snow depth species stream beds Summer browsing tall shrub habitat Timofeeva l967 tundra twigs browsed U.S. Weather Bureau Umiat area Umiat study area willow