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action potentials animal apical organ ascidian axons beat frequency behaviour Beroe Biol brain bryozoan burst Calliactis choanocytes cilia ciliary reversal ciliated groove co-ordination coelenterates colonial column comb plates comb rows conducting systems contraction ctenophores cydippid depolarization direction echinoderm Eckert ectodermal effective stroke electrical activity electrical stimulation electrode endodermal epithelial epithelial conduction epithelium evoked excitation feeding fibres flatworms function ganglion GFNN pulse Hernandez-Nicaise Horridge Hydra hyperpolarization inhibition innervation intracellular invertebrates Josephson Koopowitz larvae lophophore Lutaud Mackie marginal centres McFarlane mechanical stimulation mechanoreceptor medusae membrane Mg2+ movements Naitoh nerve-net nerve-ring nervous system neurites neurons neuropile Notoplana observations oral pacemaker Paramecium Passano pathways pedal disc Physiol physiological Pleurobrachia plexus polyclad Polyorchis polyp propagated radial receptors recorded response retraction salps sea anemone sensory cells single species specimen Spencer sponges spontaneous statocyst statolith structure studies subumbrellar surface swimming synapses TCNN tentacles tissue velocity vesicles waves zooids Zool