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9 x filipendulae alleles alleles responsible Amata Batesian mimicry black morph Bovey breeding experiments brood consisted Burgeff burnet moths camiolica chromosomes Cockayne crossed cultures dead in pupae described diapause dominant Dryja eggs ephialtes example expected Mendelian ratio female five-spotted forewing pattern forewing spots gametes genes genetics genotypic constitution giving an observed gynandromorphs heterozygous heterozygous for colour hindwing homoeosis homozygous hybrid individuals larvae Lepidoptera Linnaeus lonicerae male melanism Mendelian ratio mimicry Moreover Mullerian mimicry mutation Naumann nominal taxon normal observed ratio occur orange pairing peucedanoid morph phegea phenotype phenotypic expression populations PpRr predators prey progeny pupae purpuralis ratio of morphs recessive to red red ephialtoid red morph red peucedanoid reference Rothschild Rr°cc RRSs Ruxton Sherratt & Speed six-spotted morph specimen spots and hindwing Stauder subspecies suffused-confluent taxa transalpina trifolii W. G. Tremewan wild type wing pattern yellow morph yellow/orange YyPp Zygaena filipendulae Zygaena species Zygaenidae