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abuse ACTH adrenal glands adult rats adulthood alcohol alcohol intake alterations amphetamine amygdala anxiety anxiety disorders anxiety-related behaviors Behav behavioral tests Biagini body weight Brain Res cage Caldji Chamey compared to controls control group corticosterone cortisol dams dark phase days of age decreased depression disorders dopamine drug Early deprivation Easterling effects of early emotionality Eriksson exploratory behavior factors Fahlke female control ﬁeld ﬁnding ﬁrst ﬂeeing and freezing Francis hippocampus hole-board hormone HPA-axis Huot hypothalamus increased incubator Kalinichev Ladd locomotion locomotor activity Male Control Marmendal matemal separation Matthews Meaney modiﬁed monoamine Nemeroff neonatal matemal separation neurobiology Neurosci neurotransmitter noradrenaline nose pokes Nylander ofthe opioid peptides plasma corticosterone levels Ploj Plotsky postnatal postnatal days present study present thesis psychiatric receptors reﬂect reviewed Riiedi-Bettschen Robbins Roman room temperature separated pups separation procedure serotonin signiﬁcant differences signiﬁcantly stressors Thrivikraman voluntary ethanol intake Wigger Wistar rats Zimmerberg