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Acnistus adjusted circannual adjusted niche overlap adjusted overlap Archilochus arthropod biotic resource Brenesii Chlorostilbon circannual niche overlap CM CM CM H coexist Colibri Colwell competition Condor core species correlated Costa Rica Cuphea dimensions diurnal Ecol Ecology ecotope epiphyte Eulaema Eupherusa feeding Feinsinger flower abundance flower diversity flower numbers flowers visited food sources foraging patterns forest Gesneriaceae gross niche overlap guild effective population H CM H H H H P H Heliomaster high flower densities high-density Hamelia high-density Lobelia hummingbird species Hylocharis i_ and j_ individuals insects interspecific Kohleria Lampornis Lobelia flowers Lobelia laxiflora low-density Malvaviscus Malvaviscus arboreus Mandevilla Manettia Montane Monteverde months nectar niche breadth number of flowers overlap value plant species Plot population Fig population i_ resource base saucerottei shrub species diversity species i_ study habitats successional territorial birds territorial species traplining trees tropical tzacatl within-plant overlap