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adult elk Afognak Island American Elk antlers apparently areas bark brow tine bugling bull elk bull herd calf California cattle Cervus canadensis color cougar cow elk cow herds cow's coyotes deep snow early elk calves elk Cervus elk herds elk hunter elk population elk range elk's feeding feet forage game management grass ground growth harem master head herd bull hoofs inches incisors Jackson Hole Joe Van Wormer killed legs Leonard Lee look Manitoba mature bulls meadow migrations minutes moose mother move mule deer Murie National Elk Refuge neck normal number of points nursing Oregon plants pounds probably pronghorn reported Rocky Mountain elk Roosevelt elk rump patch rutting season seems side species spike bulls spots spring subspecies summer range teeth timber trees tule elk two-year-old usually walked wallow weather wild Wormer The World Wyoming yearling yearling cow Yellowstone National Park young bulls youngster