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acid alarm substance anal glands animals antennae ants attracted bacterium barnacles become bees beetles behaviour birds brain breeding bringing the sexes Bruce effect butterflies cabbage loopers chemoreceptors civet colony colours common chemical sense communication concentration cycle deer deermice deermouse detect developed effect egg-laying eggs feed flowers function gypsy moth hive honey-bees hormones important insects investigation Jacobson's organ known lane Burton language of smell larvae limpets lives loopers mammals mark mating membrane metres mone monkeys musk nasal nerve impulses nest nose nostrils odorous molecules olfactory clefts olfactory epithelium perfumes phero pheromone Photo by lane prey primary odours primer pheromone protozoans queen substance rabbits react reaction receptor cells reproduction salmon scent trail scent-marking secretions sense of smell sense organs sensitive sensory sexual signals snakes social species starfish stimulate strange male territory theory tion tissue turkey vultures urine visual vomeronasal organ waggle waggle dance Whitten effect workers