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activity agement aquatic areas barrier burrows canopy tree chestnut oak climate Colorado Plateau commonl cougar develop disturbance eastern hemlock ecology ecosys ecosystem editor effective experimental sign fire Florida panther forest type former NPS scientists Fossaria giant sequoia glaciers Global Change habitat hardwoods hemlock human impact lion-human located logs Meadow ment mesic mollusks monitoring Mount Rainier Mountains National Park National Park Service Natural Resources Publication noncompliant northern red oak off-trail hiking old-growth characteristics old-growth oak ozone P.O. Box Pacific Northwest Park Science Parkway pine plant pond prairie dogs preservation appeal Pulse Study regional chief scientist resource management Riper Rocky Mountain sanction sign sediments SEKI Smoky Mountains National snail species stream study plots submesic subxeric sulfate Survey terrestrial tion tional trail tree tally University upper canopy USFWS Utah vegetation virgata Washington wetlands wildlife Yellowstone Zion Canyon Zion National Park