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adult survivorship average dates average fruit removal avian seed dispersal background color Barkley Sound bird abundances black fruits bracteosum bracts British Columbia census interval character displacement competitive interactions dates of fruit environmental conditions evaluated evidence for character foliage coloration forest understory habitat frugivores fruit availability distributions fruit color fruit displays fruit maturation dates fruit persistence fruit production fruit removal rates fruit-eating birds fruiting distributions fruiting patterns fruiting phenologies fruiting season habitat-matching immigration island area island distributional patterns island size classes Julian Date large islands light and soil Lonicera involucrata northwestern crow null model analyses number of fruits number of seeds parviflorus patterns in fruit phenological overlap phenological patterns population densities PRNP proportion of fruits racemosa red fruits resource axis Rhamnus purshiana Ribes Ribes divaricatum Rubus parviflorus Rubus spectabilis Sambucus racemosa sapling seeds trapped shrub species small islands total fruiting frequencies total number trial Vancouver Island western hemlock Willson