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ability altered angle animal animal's antennae appear arthropods attracted beam bees beetle behaviour Biol body caterpillars cells changes chemical chemoreceptors colour compound eye courtship crabs crawl crustacea dark detected direction dorsal effect example female flies foragers frequency function gradient gravity ground hairs head heat hive honeybees horizontal humidity illuminated increased insects intensity invertebrates larvae legs lower male moths move movement nerve nervous system nest normal objects occur ocelli odour olfaction ommatidia ommatidium orientation Orthoptera Paramecium pattern perceived phototactic Physiol physiological pigment placed planarian polarized light position preference pressure prey produced rays react reaction receptors response scent screen seems sense organs sensitive sensory side similar skatol sodium chloride sound species speed spiders statocysts statolith stimulation stripes substances sucrose surface swim temperature threshold ticks tube turn tympanal organs Uber vergl vertical vibrations visual wasp wave wavelengths woodlice Zool