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Adult Females age classes Aggressive Adult Males Aggressive Females Aggressive Males appeared artificial food Bears Marked bluffing breeding season brown bear cent Class Specific Designation CLASSIFICATION COUNTS classified CM CM concentration sites copulation Craighead Creek-Hayden Valley area Cutlip different males direct count displayed dominance classes dominance structure dominant animal Dominant Male estrus period exhibited family groups females in estrus Females Observed fighting grizzly bears grizzly population hierarchy identified females July June June 26 Kodiak Island large number lesser individuals lodgepole pine Marked Bears Montana Montana State University natural food Nights Trap Number 12 NUMBER OF OBSERVATIONS Number XX obtained Offspring Weaned order of dominance orphaned Park-wide population position presented in Table reproductive behavior Scarface Schnabel Method single female social rank social structure Storer and Tevis Sub-Dominant Male succinylcholine tion Trout Creek Trout Creek-Hayden Valley two-year olds Weaned Yearlings Yellowstone National Park Young Adults