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autecology available light available water capacity Black raspberry blackberry choke cherry compositional gradient continuum Crataegus Curtis and Mcintosh Curtis l959 decimal segment decrease deep shade conditions driftless area dry forest dry-mesic forest elderberry environmental conditions environmental factors feet per acre Figure l0 forest communities forest group Goff gray dogwood inches at DBH inches in height increase Manitoba maximum diameter mesic deep-shade forest mesic medium-shade forest mesic-oak forest milacres per milacre modal overstory basal area overstory species peak importance plots poison ivy Quebec Quebec and Minnesota reaches a maximum reaches its optimum reaches its peak red raspberry Ribes missouriense Rosaceae Rubus idaeus sampling seedling density shrub cover shrub density shrub importance values shrub layer shrub species soil horizons soil texture southern Wisconsin upland species reach stem diameters study area sugar maple total importance total shrub importance upland forest gradient Wisconsin upland forest Witch hazel woodbine xeric xeric conditions