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Adult Fish algal groups benthic benthos biological biotic populations Blue-green Algae buffering carbohydrate carbon coefficient of variation compartment turnover components correlation daily pool sizes degree of recycling detrital compartments detrital pools Detritus Settled Diatoms differences in turnover dissolved inorganic nutrients dissolved organic matter ecosys ecosystem model fast pools flux rates function gain rate gm dry Green Algae growing season hypothesis individual compartments inorganic nutrient pools inputs Koonce labile detritus Lake Texoma Lake Wingra ecosystem longer turnover loss rate nitrogen and phosphorus nitrogen cycle Nitrogen Pools nonliving nutrient dynamics Odum Organic Matter Pools pelagic zone phosphorus pools plankton Pomeroy postulated rates and turnover recycling refractory detritus result similar simulation size and flux sizes and flux slow pools spring and summer stable Standard Deviation system compartments system flux Table temporal variation total organic matter total system pool turn uptake variability in turnover various compartments WINGRA III model zooplankton