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40-hectare study area abundant vertebrate acorns Arrowsic Average number Back River basal area Bill Holt Black-throated Green Warbler canopy control cm DBH Common Yellowthroat control areas control blocks Department of Wildlife eastern white pine fence line ﬁeld Figure ﬁrst ﬁve forest canopy gaps forest stands gaps created goal graduate students growing on mesic habitat han/est harvest blocks harvest conducted harvest gaps harvest-created canopy gaps harvested areas Holt Forest Holt Research Forest Holt Woodlands Research Kennebec River land ledge gaps located long-term research Maine’s monitor natural number of salamanders number of stems oak-pine forest ecosystem partial harvest Photo plants red maple red oak redback salamanders region research program salamanders found seed seedling and sapling Sewall Pond small mammals south farm subquadrats Tammy Clark timber harvest timber management tion trailing arbutus tree gaps University of Maine white pine white-footed mice White-throated Sparrow Wildlife Ecology Winter Wren Witham Woodlands Research Foundation xeric