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a a a a airway closure alveolar Appl arterial oxygenation blood flow cardiac output central blood chest chronic obstructive pulmonary closing volume CN CN CN COPD Subjects decrease diaphragm diffusing capacity diving dyspnea effects of immersion effects of prone emphysema exercise in patients expiratory flow gas exchange groups head-out immersion heart rate higher hydrostatic balance hydrostatic pressure hypoxemia I I I I I I I I Increased Oxygen Uptake inspiratory muscles L/MIN lactate lower maximal mechanics MEFV Metabolic mmHg normal and COPD normal subjects obstructive pulmonary disease patients with chronic patients with COPD perfusion Physiol Physiology postural changes prone immersion Pulmonary diffusing capacity reported Resp respiratory rest static lung load static lung volumes stroke volume submaximal supine immersion supine position swimming thorax tidal volume unchanged University of Wisconsin-Madison upright immersion values venous