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abundance activity additional analysis analyzed animals backcountry basin beach behavior Bighorn birds brook trout Cave Research Foundation collected Colorado River completed conodont continued coyotes Creek cutthroat trout Death Valley deer density determine distribution dune Ecology ecosystem fauna field fire fish forest Geological Glacier Grand Canyon Gross Acreage Guadalupe Mountains habitat HISTORICAL SITE Visitors impact Indiana bats indicate Island Lake levels located mammals Mammoth Cave meadow monitoring Mountains National Park N-l N-l NATIONAL HISTORICAL NATIONAL MONUMENT Visitors NATIONAL PARK INVESTIGATOR National Park Service nesting observed occurred PARK Visitors planned plant plots population present range recreation reproductive rock rodents samples sand scat season sediments small mammal soil species specimens spring study area SUMMARY OF PROGRESS summer survey temperature trail transects trapping trees trout U.S. Geological Survey University SUMMARY vegetation Wildlife winter Yellowstone Yellowstone National Park