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abundance activity adaptation adult analysis animals Apodemus araneus autumn bank vole bats behaviour Berchtesgaden National Park Biology body weight breeding canids carnivores changes Clethrionomys composition conservation correlated culpeo decrease Department of Zoology desert dispersal distribution diversity dominant Dorcas gazelle dynamics ecological ecosystem endemic environment environmental Eocene extinction factors fauna females foraging forest foxes frequency gazelle genetic geographic glands groups growth habitat hibernation home ranges hunting increase individuals Institute islands juvenile litter males mammalian mammals mating morphological mountain National Park natural North observed occur Ochotona Oligocene Paleocene patterns period Pleistocene population density predation present prey pups raccoon dog region relatively reproductive Research rodents roe deer season sexual shrews small mammals social Soricidae species structure summer survival temperature territories total number ungulate University variation vegetation Vulpes whales wild wildlife winter Zool