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adaptability advanced-generation alleles breeder breeding program Buijtenen characteristics chromosomes clonal clones combining ability Conf conifers crosses desired developed diallel disease Douglas fir economic effects environmental established Eucalyptus example exotic fertilization Figure Forest Genetics forest management forest tree improvement forestry fusiform rust gene complexes gene conservation genetic base genetic gain genetic tests genetic variation genotypes grafts grow growth hardwoods heritability hybrid important inbreeding individual trees insects intensive interaction IUFRO Jour land race loblolly pine method Namkoong natural stands needed North Carolina obtained occur operational parents phenotype Pinus plantations planting pollen polyploid potential Proc progeny test propagules provenance radiata regeneration Research result root seed orchard seed production areas seed source seedlings slash pine southern pines species specific gravity taeda tracheid traits Tree Impr tree improvement program tropical types U.S. Forest Service usually variability variance vegetative propagation wood properties wood qualities wood specific gravity Zobel