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abdomen adapted Adult beetles amateur Coleopterist animal antennae appear appendages aquatic arrow shows obscure attack Augomela hypochalcea Australian beetles AUSTRALIAN ENTOMOLOGICAL Balanophorus BEETLE BIOLOGY Beetle larvae beetle pupae beetles possess book on Australian borers Buprestidae Campodeiform Carabidae Cerambycidae Cetoniinae Chrysomelidae Cleridae Coccinellidae cockroaches Coleoptera Cucujidae Curculionidae cuticle Dermestidae and Ptinidae Dytiscidae eggs elytra elytron Erotylidae Eruciform families are entirely families e.g. family Scarabaeidae fascicles feeding habits female five-segmented flight fore wings four-segmented frequently Gyrinidae harden and darken heavily armoured heteromerous hind wings hindbody Hydrophilidae inquilines insects known labrum larvae and adults larvae Fig larval stages latter Leaf beetles legs Leptospermum Lucanidae male mandibles Melyridae mesothorax metathorax mouthparts non-Australian normally Numerous palpi palpus Passalidae pentamerous plant stems plant-feeders predators pseudo-tetramerous arrow shows pseudo-trimerous Rhipiphoridae Rose chafers Scarabaeidae sclerotisation Silphidae soft and pale SOUTH-EASTERN AUSTRALIA specialised species Staphylinidae and Histeridae Syndesus cornutus Tarsal formulae tarsus Tenebrionidae thorax underwings wasps Weevil