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4th hind toe A.A. Echelle Adult male Anolis amphibians Andre's and Providencia anoles Anolis cupreus Anolis villai showing average 169 basal CARIBBEAN NICARAGUA Central American mainland chocolate-colored closely related concolor continental shelf Corn Island Costa Rica counts dark with narrow darker dewlap dark dorsal crest Dorsal ground color extending forward females fourth hind toe frontal concavity gravid H.S. Fitch head length heavily keeled Holotype IGUANIDAE interparietal Iris dark Jaime Villa large supralabials level of groin lizards loreal rows Loveridge 1929 lowered sea level males average 51.5 markedly larger mid-body middorsal Midventral scales Milwaukee Public Museum pale rim paler Photo by R.W. pinchoti population Providencia Islands R.W. Hender rodriguezi San Andre's San Jose Scale rows scales on top sericeus series of scales sexual size difference species stage of lowered supraorbital semicircles separated surface dark sympatric tip of snout UKNHM University of Kansas Ventral scales Ventral surface dusky Vertebrate Division widened lamellae