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adaptation adult animals Behav bulbectomized change sex chemical components of fitness conspecifics cycle days of age dead prey discriminate dish dominant effect Emy's environment environmental estradiol estradiol benzoate estrogen estrous estrous cycling fish friendly approach behavior gland gonadal greeting H-Y antigen hamsters havior hermaphroditic hormonal human companion hysterectomized hysterectomy increase individual initiated injection intranasal juveniles kittens lactating latencies learning licking litter male rats male removal maternal behavior mhrm mhrn milk mother nest nonpregnant females occur odors olfaction olfactory onset of maternal ovariectomized females parturition period phenotypic pheromone phylogenesis Physiol play population postpartum pregnancy preputial gland primary referent progesterone prolactin protogynous pup exposure pup stimulation purring quick/noisy Reinboth retrieving rubbing S15 Rennie secretion sensitization latencies sex ratio sex reversal sexual Shapiro Siegel and Rosenblatt slow/quiet social species squamipinnis studies subgroup suckling tertiary referents tion urine uttered vaginal vocalization week zinc sulfate