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ammonites Angola Annals anterior arch archosaurs articulates base become BMNH body chamber Broom Campanian Cape caudal vertebrae closely Cobban coiling collections Collignon complete compressed Coniacian consists constrictions Cooper Cretaceous cylindraceum described Description diameter differs Diplomoceras Discussion distal dorsal early edge elements Erythrosuchus africanus Eubostrychoceras example fauna Figure flanks fossils fragments genus Geological Gower History holotype Howarth incomplete Kennedy Klinger known KwaZulu Lateral view LIBRARY locality lower Maastrichtian Madagascar material Matsumoto maxilla medial Middle narrow Natural occurrence original ornament Paleontology phragmocone position possibly posterior present preserved probably proximal referred represented ribs rounded Santonian shaft shell shows similar single skull slightly Society South African Museum Spath species specimens St Lucia Formation suggests surface suture text-fig tubercles Turonian Type species umbilical Upper Upper Cretaceous venter ventral ventrolateral vertebrae whorl whorl section