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activity adult females adult males age-class Alaska Alaska Peninsula animals average backcountry baits Banff National Park bear damage bear habitat bear management bear Ursus arctos bears killed behavior biology and management black bear population black bears Ursus breeding brown bears captured cementum conifer contact index Craighead critical habitat dens density Dept determined distribution elevations Erickson estimated feeding Fish and Game Fish and Wildlife forest grizzly bears habitat habitat types harvest Hedmark home range hunters hunting increased indicated Island IUCN IUCN Publ Japanese Jonkel and Cowan Karluk Kodiak LiCl litter located Mammal Martinka Montana mortality movements number of bears observed occurred percent period Poelker polar bears predation premolar protein reported reproductive sample scats season sex ratio skull snow species study area subadult surplus killing Table tion trees U.S. Fish U.S. Forest Service Univ vegetation Wildl Wildlife Service winter yearlings Yellowstone National Park