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absorption abundance estimate acoustic axis acoustic cross-section acoustic data acoustic instruments acoustic surveys amplitude applied average bandwidth beam pattern beam width behaviour bubble cage calculated calibration coefficient considered cruise track curve decibels depends depth described detected determine direction distance Dmax dual-beam echo counting echo energy echo integration echogram EDSU Equation equivalent beam angle error example explosion factor fish density fish distribution fish migrate fishery frequency hydrophone insonified intensity length lobes MacLennan mean method noise observed pelagic trawl physostomes plankton pressure problem propagation proportional pulse duration random randomly range received signal region resonance samples scattering schools scientific echosounder sea-bed sensitivity ship side-scan sonar sonar sound speed sound waves species sphere standard target Suppose surface surveyed area swimbladder target strength technique thermocline threshold tilt angle towed transducer transects transmission transmitted trawl tungsten carbide TVG function variance vertical vessel volume wavelength