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25 cm dbh AGESQ average background vegetation beech black cherry canopy gap canopy trees CM H CM rH cucumber tree decreased density disturbance regime elevation estimates expanded gap fraction of gap fraction of land gap age gap closure gap phase gapmakers H CM H rH hardwoods Hueston Woods Joyce Kilmer Kalanu Prong large gaps lateral extension growth least 25 cm maximum mesic Mount Guyot northern red oak point-centered quarter method RA1SQ rate of gap red maple regression relationship relative density relative importance rH CM rH H sampled sapling height growth significant silverbell slope small gaps southern Appalachians species composition species diversity stems striped maple structural diversity Study Area sugar maple Table Tionesta tolerant species transect trees at least tulip tree understory Walker's Cove white ash white basswood Wisconsin importance values yellow birch yellow buckeye