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aalge adult breeding adult non-breeding Advertising-call Africa Alarm-call Alcidae Arctic Tern Behaviour bill birds birds ringed Birkhead Borodulina breeding season brood buff chicks Clupeidae clutches coast colony Common Tern copulation Coquet Island courtship crown crustaceans darker Dementiev and Gladkov Downy Young dusky edge eggs especially feathers feeding fish fledging flight flight-feathers flocks forehead Goodwin ground hatching head Heinroth hindneck hirundo Hulsman I C T Nisbet immature incubation inner primaries Island juvenile Kandalaksha Bay Kotov Laridae less Little Tern mainly mantle mate moult Murton nape neck nest nest-site nominate numbers outer primaries pairs pale grey Palearctic paler parents plumage Post-breeding primary coverts primary moult RMNH roost rump Sandwich Tern scapulars sexes shorter similar slightly sometimes Spitsbergen Sterna Sternidae tail tail-feathers tarsus Tern tertials tinge underparts upper wing-coverts upperparts USSR usually Voice West Germany west Palearctic wing winter