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THE GREAT TIT IN wyTHAM MoRTALITY AT DIFFERENT
THE coAL TIT IN PINE PLANTATIONs 8o_
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adaptive adult ratio Andrewartha and Birch annual average clutch average number birds Blackbird Blue Tits Botanic Garden bred breeding pairs breeding population breeding season broad-leaved woods brood-size caterpillars cent chick clutch-size Coal Tit cocks Collared Flycatcher colonies correlated date of laying decrease density-dependent earlier early eggs evidence evolved factors favourable feeding females figures fledged fluctuations food shortage food supply Forest of Dean habitat hatched hectares Hence hens higher increase individuals insects juvenile Kittiwake Kluijver larvae later left the nest males Manx Shearwater Marley wood mortality natural selection nestlings normally number of breeding number of young parents period Perrins Pied Flycatcher pine population density predators presumably probably proportion Ptarmigan Quelea Red Grouse reproductive rate Schiiz Scots pine second broods seeds Short-tailed Shearwater Song Thrush species studied suggest summer survival Table Tawny Owl territorial behaviour unusually variations weather White Stork winter Wood Pigeons Wynne-Edwards Wytham yearlings Yellow-eyed Penguin young raised