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alfalfa CM CM CM CM CM H CM H CM CM OO CM rH concentration considerable content in oat content of nitrogen correlation deficiency symptoms dry matter element factors four replicates Goodall and Gregory grain yield green growth growth medium H CM CM H H H H H rH harvest heading stage HHHH increase inflorescence jointing stage Jointing to milk leaf fractions lower leaves lower plant middle leaves milk stage Milligrams Mount Pleasant samples nitrate nitrogen fertilization nitrogen supply nutrient oat leaves occurred ONCO ooco OOOO OOOO percent of dry percentage content phosphorus and potassium phosphorus content phosphorus supply plant analyses plant sap plants vere plots potassium content rapid tissue tests relationship rH H second sampling senescence soil stages of development stem third sampling tillering top leaves total content total nitrogen translocation treatments trogen uptake varied whole plant yellowing