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active space aggressive behavior androgen animal's anosmia anosmic areas scented attracted Behav Biol Bronson Carr castrated castrated male chemical communication chemical signals clean box clean cages conspecific odors Doty Drickamer effect encounters estrus cycle evidence Ewer female cages female flank marking female urine flank gland flank glandectomized male flank marking rates function gland secretion glandectomized male cages glandectomized male scented golden hamster home cage home range hormonal house mice Johnston Jones and Nowell latency to enter Lindzey male and female male odors male scented areas male urine mammals maniculatus mating behavior median Meriones unguiculatus Mesocricetus auratus Mongolian gerbils Mugford and Nowell Mykytowycz nonestrus nonreward normal male cages normal male scented odor conditions odor sources odors associated olfaction olfactory open field Pheremones Physiol preference tests preputial gland reproductive rodents scent marking sebum skin glands source of odors species stimulus box stressed studies Thiessen tunnel vaginal discharge vaginal marking Vandenbergh