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A-male abyssinica activity Ad-unit Addis adolescent adult females adult males Afincha all-male group alpha female animals approached Araki Arat Ayin baby behaviour biomass Bote C-group C-R bond calling and replying chased Chereka cliff copulation core moving range DH-unit distribution Djera dry season Dunbar E-herd E-male group Emietgogo female cluster Festuca fighting food intake frequency friendly gelada baboons Gich Plateau Gira GK-unit Gojira grass blades grooming interactions grooming relations grooming relationships hamadryas baboons Haya herding behaviour Hy-unit individual distances infant Japanese monkeys junior free-lances juvenile Kadadit Kegin Kokebu Kz-unit leader male lip-roll Lobelia loud defensive sound Meji Meskel Mesu-A mounted movements multi-herd natal unit Nora observation period oestrous female one-male units Papio plants population presenting primates puberty males rainy season second male Sidist Simen social spatial species steppe Tadji Tara threatened Tn-females unit members vegetation vocal Wakame