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100 females access roads active age classes Ambrosia dumosa animals annual approximately Arden Study Area Arizona Beaver Dam behavior Berry Brattstrom Bureau of Land captive tortoises Copeia County creosote bush Department of Fish desert mallow Desert Tortoise Council Desert Tortoise Natural desert tortoise populations Desert Tortoise Relocation Fish and Game forage frequency Gopherus agassizi grass growth rate hatchlings home range impact Joshua Tree juveniles Land Management located Luckenbach males to 100 marked methods Miller Mojave Desert Mountains Museum Nevada Nov Nov off-road vehicle P.O. Box Patterson percentage period Population density recapture reptiles San Bernardino County San Diego scats scutes season sex ratios sexual maturity shell shrub small tortoises Southern California Edison Sphaeralcea ambigua square miles Stephanomeria subadults survey temperatures toises tortoise habitat Tortoise Natural Area Tortoise Relocation Project transmission lines Turtle and Tortoise Utah Utah population Vegas vegetation vocalizations wild wildlife winter Woodbury and Hardy