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abundant Alan Nelson American Bison Society antlers autumn balsamroot battles beasts bighorn sheep birds blue grouse bucks buffalo bulls buffalo calves buffalo herds buffalo refuge bull elk burrowing Canada Charles Allard climb coat Columbian ground squirrel coyote Danny deep snow dominance early environment extinction fat reserves fawns feed feet fenced Flathead Valley forage forest geologic grasses and forbs grassland habitat hair hide hunters higher elevations hooves horns humps hunting Jon Cates large grazing animals leaves M.J. Gordon mating season Michael Sample Michel Pablo's migrate south miles moisture Montana Historical Society mountain goats mule deer National Bison Range nest North American continent north slopes number of buffalo Palouse prairie Plains Indians pounds predators pronghorns roamed rock Rocky Mountain rodents rutting season seeds Short-eared owls small mammals soil species spring summer sweeping tail thick thousands U.S. Fish Voles weigh Whitetails wildflowers William Hornaday winds winter woodlands yellow-bellied marmot young