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Acanthuridae acoustical energy adult parrotfishes amount of calcareous Bahamas Bardach basic benthic Bimini calcareous material removed CM CM CM coelestinus coral reef coral substrate CRUNCH sounds differences in feeding feeding activities feeding behavior feeding fishes feeding habit feeding sounds recorded frequency range frequency structure g of calcareous grazing parrotfishes H 4J habitat hard substrates herbivorous Heron Island hr on July Hydro-Lab hydrophones July 23 Fig July 24 kg/ha/yr Marine bio-acoustics monitoring number of feeding occur parrotfish feeding sounds parrotfish grazing passive acoustical technique R. G. Busnel Randall rasping reef fishes removed per CRUNCH rH H saturation dive Scarus guacamaia SCRAPE and CRUNCH SCRAPE sounds sexual dichromatism small parrotfishes sonic sonograms Sparisoma viride species of corals species of parrotfishes spectral analyses stridulation surgeonfishes Tektite temporal spacing Texas A&M University Travolga types of feeding underwater underwater acoustics underwater habitat visual observations