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adult and subadult adult central females adult central males adult females adult floating females adult males adult nucleus female adult peripheral males affinitive Arashiyama West troop Attentive/locating behavior factors behavior units Bravado centroids Confident aggressive criteria difference in behaviors direct discriminant functions factor analysis females subadult havior individual Infant comfort seek Infant wean Insecure aggressive interactions ITANI Japanese macaques juvenile central females juvenile central males juvenile females juvenile floating females juvenile floating males juvenile males lambda criterion Locomotor play lowest Macaca fuscata male behavior Male courtship Male sex males score highest Mother wean Mother-infant conflict multiple discriminant analysis null hypothesis offspring peripheral behavior peripheral females primatologists Protective/defense Reassurance/affinitive Receive aggressive play significant difference spatial status subadult central females subadult females subadult floating females subadult peripheral males Submissive troop members Troop of Japanese twelve role groups uterine group variables Wean sounds YAMADA