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1980 symposium acre aggregation pheromones amphibians annual antiaggregation Arizona attack Auffenberg baited bark beetle Beaver Dam Slope behavior Berry Bureau of Land Burge burrows Bury California captive tortoises Code County Coombs data base Dendroctonus densities Department of Agriculture desert tor Desert Tortoise Council desert tortoise Gopherus ecology eggs Fish and Wildlife forage Forest Service forest type gopher tortoises Gopherus polyphemus grazing growth habitat Hashagen hatchlings Herpetologica Interior International Turtle inventory ipsdienol Lake Havasu City Land Management Lodgepole pine Long Beach Luckenbach 1982 males March Mojave Desert mortality mountain pine beetle Nevada Patterson percent pheromones plot predators Proceedings reptiles Reptilia Riverside Rosskopf sample scutes semiochemicals shell softwood types southern species spruce beetle timberland tion toise tortoise Gopherus agassizii tortoise populations Tortoise Society Journal traps Trotter Turner Turtle and Tortoise U.S. Department Utah Valley Vegas verbenone Western Wildlife Service Woodbury and Hardy WWDB