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abundance Acacia adult cheetah adult female adult groups adult males aggression appears approach attack behavior birth bite Bourliere breeding captive cheetahs carcass cats chase cheetah cubs cheetah groups cheetah kills cheetah prey cheetahs chased comm courtship dew claw distances East Africa estrus estrus female exhibited feces feeding Felidae fighting Flehmen forepaw four cubs frequently Graham and Parker Grant's gazelle grassland habitat herd Herdman hunting success impala individuals jackals Kenya Kierney kongoni Kruger Park Kruuk leopard Leyhausen Lion Country Safari litter markings mating meat months old mortality mother Nairobi National Park Nairobi Park neck observed Photo population pounds predators predatory prey animals prey species prey's probably R. L. Eaton reedbuck response San Diego savanna Schaller Serengeti sex ratio Shortridge smelling social South West Africa stalking Stevenson-Hamilton territory Thomson's gazelle ungulates urine urine-mark warthogs waterbuck wild cheetahs Wildebeest World Wildlife Safari yards young zoos