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absorption acclimation action spectrum adaptation algae aquatic Biol biological Bjorkman boundary layer C4 plants canopy cells changes chlorophyll chlorophyll content chloroplast chromosome CO2 pressure coefficient crop diffuse dose ecological effects energy environment environmental enzymes factors far-red light fire flowering fluence rate function germination grown habitats heat tolerance high light higher plants Ichikawa increase inhibition irradiance Larcher leaf area leaf temperature light intensity light levels light regime lipids low temperature measurements membrane mesophyll metabolism mutation natural Oecologia ozone photoinhibition photon photoperiod photosynthetic rate photosynthetically active radiation physiological phytochrome phytoplankton pigment plant growth Plant Physiol protein quantum flux densities quantum yield radiosensitivity ratio reaction resistance response RuBP RuBP carboxylase seedlings seeds sensitivity shade plants shadelight Sinapis alba Smith soil Solanum dulcamara solar UV somatic species spectral spectrum stress studies sun and shade surface thermal thermophilic tion tissue Tradescantia UV-B vegetation wavelengths