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Abies balsamea Alnus alternate isolation animal borne propagules area and isolation average slope bedrock birds Canada caribou Connor and McCoy Cornus canadensis density of propagules Dept distance and isolation dry-mesic Ecology effective isolation habitat habitat diversity Halberd higher immigration rates higher plant species inversely island area Island slope island species number islands and mainland islands in Lake isolation distances Lake Kelvin Lake Nipigon large islands loge species number lower extinction rates lower immigration rates MacArthur and Wilson mainland exposure mesic moisture type moss Nipigon Island Biogeography Nipigon River number of species NW Ontario Ontario Phytol Picea plant species diversity populations Populus balsamifera predictor of island Preston probable dispersal mechanisms random walk recipient island ruffed grouse Sample islands sampling effect seeds Simberloff simple distance small islands Sorbus decora source area species on Lake stepping stone suggests lower immigration t-test Table topography type variables Viola nephrophylla wet-mesic