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" Echinulation occurred at times in all of the species studied. In some cases part of the spores from a colony were echinulate while others were smooth. "
American Journal of Botany - Page 461
1917
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Taxonomic Characters of the Genera Alternaria and Macrosporium

John Asbury Elliot - Alternaria - 1916 - 56 pages
...relatively short time—two to six weeks—in some species there is a marked secondary development. is variation in shape within a single species, but...species. Generally spores darken with age. Alternaria dianlhi and A. brassicae var. microspora grown at IO° showed marked color change, both being very...
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